Annabel Hayes--born, baptized, and orphaned in the sleepy conservative town of Dayton, Tennessee--is thrilled to find herself falling quickly and deeply in love with George Craig, a sophisticated attorney newly arrived from Knoxville. But before the end of their first year of marriage, their lives are beset by losses. The strain on their relationship is only intensified when John T. Scopes is arrested for teaching Charles Darwin's theory of evolution at the local high school.

Foreshadowing today's culture wars, the trial against Scopes is a spectacle unlike any the country has seen. William Jennings Bryan--a revered Southern politician--joins the prosecution, pitting himself and his faith against the renowned defense attorney Clarence Darrow. Journalists descend in a frenzy, thrusting the town and its citizens into the national spotlight. And when George joins the team defending Scopes, Annabel begins to question both her beliefs and her vows.

As the ongoing trial divides neighbor against neighbor, it also divides the Craigs in unexpected ways. But in the midst of these conflicts--one waged in an open courtroom, the other behind closed doors--Annabel will discover that the path to her own evolution begins with the courage to think for herself.